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California Standards Biology


Standard 8f Preknowledge

8f) ***Students know how to use comparative embryology, DNA or protein sequence comparisons, and other independent sources of data to create a branching diagram (cladogram) that shows probable evolutionary relationships.

CALIFORNIA FRAMEWORKS SUMMARY:

The area of study that connects biological diversity to phylogeny, or the evolutionary history of a species, is called systematics. Systematic classification is based on the degree of similarity between species. Thus, comparisons of embryology, anatomy, proteins, and DNA are used to establish the extent of similarities. Embryological studies reveal that ontogeny, development of the embryo, provides clues to phylogeny. In contrast to the old assertion that “ontogeny recapitulates phylogeny” (i.e., that it replays the entire evolutionary history of a species), new findings indicate that structures, such as gill pouches, that appear during embryonic development
but are less obvious in many adult life forms may establish homologies between species (similarities attributable to a common origin). These homologies are evidence of common ancestry. Likewise, homologous anatomical structures, such as the forelimbs of humans, cats, whales, and bats, are also evidence of a common ancestor. Similarity between species can be evaluated at the molecular level by comparing the amino acid sequences of proteins or the nucleotide sequences of DNA strands. DNA-DNA hybridization, restriction mapping, and DNA sequencing are powerful new tools in systematics.

Approaches for using comparison information to classify organisms on the basis of evolutionary history differ greatly. Cladistics uses a branching pattern, or cladogram, based on shared derived characteristics to map the sequence of evolutionary change. The cladogram is a dichotomous tree that branches to separate those species that share a derived characteristic, such as hair or fur, from those species that lack the characteristic. Each new branch of the cladogram helps to establish a sequence of evolutionary history; however, the extent of divergence between species is unclear from the sequence alone.

Phenetics classifies species entirely on the basis of measurable similarities and differences with no attempt to sort homology from analogy. In recent years phenetic studies have been helped by the use of computer programs to compare species automatically
across large numbers of traits. Striking a balance between these two approaches to classification has often involved subjective judgments in the final decision of taxonomic placement. Students can study examples of cladograms and create new ones to understand how a sequence of evolutionary change based on shared derived characteristics is developed.